A ‘model-to-radiance’ comparison of simulated brightness temperatures from the Hadley Centre Global Environmental Model 2 with measurements from the High Resolution Infrared Radiation Sounder/4 (HIRS/4) instrument onboard the MetOp-A satellite is presented. For the all-sky, the model overestimates brightness temperatures in the atmospheric window region with the greatest biases over areas associated with deep convective cloud. In contrast to many global climate models, much smaller clearsky biases are found indicating that model clouds are the dominating source of error. Simulated values in upper atmospheric CO2 channels approximate observations better as a result of compensating cold biases at the poles and warm biases at lower latitudes, due to a poor representation of the Brewer Dobson circulation in the 38 level ‘low-top’ configuration of the model. Simulated all and clear-sky outgoing longwave radiation (OLR) evaluated against the Clouds and the Earth’s Radiant Energy System (CERES) and HIRS OLR products reveal good agreement, in part due to cancellation of positive and negative biases. Through physical arguments relating to the spectral energy balance within a cloud, it is suggested that broadband agreement could be the result of a balance between positive window biases and unseen negative biases originating from the water vapour rotational band in the far infrared (not sampled by HIRS).